Mother Miranda is the main antagonist of Resident Evil Village.
Mother Miranda is a scientist who was born in a Eastern European village in the late 19th century who lost her daughter Eva during the Spanish flu pandemic. While entering a cave with the intent of following her daughter in death, a despondent Miranda is infected while coming across a fungal superorganism called the "Megamycete" within the cave that granted her immortality, the ability to assume the form of others, and the collective knowledge of those the fungus came in contact with prior.
Miranda uses her gained abilities to experiment on the villagers for years, creating the Cadou parasites in failed attempts to create an ideal vessel to revive Eva in. Miranda is later revealed to be the mentor of Umbrella Corporation founder Oswell E. Spencer and inspired him to achieve his vision of human evolution, making Miranda the indirect instigator of all the major events that transpire in the series. She also provided her assistance to the Connections by providing them the means to create Eveline, later learning of Rosemary Winters from them as she abducts the infant so she can resurrect her daughter. She is ultimately killed by Ethan Winters who rescues Rosemary from her before sacrificing himself to destroy the Megamycete.